International Journal of Biodiversity and Conservation 2141-243X Academic Journals 10.5897/IJBC2022.1571 Takeda Shingo Murayama Takehiko Nishikizawa Shigeo Nagaoka Atsushi * E-mail: shingotakeda@hotmail.co.jp 2023 31 01 2023 31 10 2022 23 01 2023 15 1 13 29 This is an open-access article distributed under the terms of the Creative Commons Attribution License 4.0, which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. This article is available from http://politicalwaffle.uk/journal/IJBC/article-abstract/B459FC870326 The full text article is available as a PDF file from http://politicalwaffle.uk/journal/IJBC/article-full-text-pdf/B459FC870326 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51